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College of Life Sciences
Biology
John Kauwe

John "Keoni" Kauwe

University President, BYU–Hawaii and Professor, BYU
Curriculum Vitae
Biology
Website: https://kauwelab.byu.edu
Email: kauwe@byu.edu

4102 LSB - Brigham Young University
Provo, UT 84602

801-422-2582

Biography

Dr. Kauwe is an internationally recognized researcher specializing in Alzheimer’s disease genetics. He earned his Bachelor’s and Master’s degrees at BYU Provo. He then completed his Ph.D. at Washington University in St. Louis in 2007 and a postdoctoral fellowship at the Washington University School of Medicine in 2008. He joined the faculty at BYU in 2009 and is a Professor in the Department of Biology. He served as chair of the department in 2018-2019 and dean of graduate studies in 2019-2020. Currently, he is the president of Brigham Young University–Hawaii.

Dr. Kauwe has over 120 peer-reviewed publications and his research is funded by more than $6 million dollars in extramural grants. He has made important contributions towards discovering more than a dozen new genetic risk factors for Alzheimer’s disease. His work has been recognized nationally and internationally as evidenced by his appointment as a Senior Editor for the journal Alzheimer’s & Dementia, his recent service as a panelist for the 2015 National Institutes of Health Summit on Alzheimer’s disease Research, and as the scientific lead for the international Alzheimer’s disease DREAM Challenge.

Research Interests

Dr. Kauwe's research leverages novel phenotypes and approaches to characterize the genetic architecture of Alzheimer’s disease with the ultimate goal of finding a cure.

Research in the lab also includes studies on prevention, diagnosis, and the genetic basis of rheumatic heart disease and the use of genomics to address evolutionary questions in several species of fish and reptiles.

Teaching Interests

Within the Department of Biology, Dr. Kauwe has taught Biology 130, Introduction to Biology and Biology 463, Genetics of Human Disease. He is also committed to providing high quality mentored research experiences in his lab. His research is designed to include significant contributions from BYU undergraduates. His students have authored and co-authored more than three dozen manuscripts and received many awards for their research efforts.

Honors and Awards

  • 2015 BYU Sponsored Research Award, Brigham Young University (2015 - 2015)

Professional Citizenship

  • Reviewer, Ad Hoc Reviewer, BMC Bioinformatics (2012 - Present)

Courses Taught

Publications

Brandon D Pickett Elizabeth M Wallace Perry Gene Ridge John Sai Keong Kauwe C L Sanders G B Rattinger M S DeBerard A G Hammond H Wengreen John Sai Keong Kauwe M Buhusi J T Tschanz MTW Ebbert T D Jensen K Jansen-West J P Sens J S Reddy Perry Gene Ridge John Sai Keong Kauwe V Belzil L Pregent M M Carrasquillo D Keene E Larson P Crane Y W Asmann N Ertekin-Taner S G Younkin O A Ross R Rademakers L Petrucelli J D Fryer S M Wright S L Jensen K L Cockriel B Davis J T Tschanz R G Munger C D Corcoran John Sai Keong Kauwe L A Cannon-Albright N L Foster K Schliep J M Farnham C C Teerlink H Kaddas J Tschanz C Corcoran John Sai Keong Kauwe D Patel J Mez B N Vardarajan L Staley J Chung X Zhang J J Farrell M J Rynkiewicz L A Cannon-Albright C C Teerlink J Stevens C Corcoran J D Gonzalez Murcia O L Lopez R Mayeux J L Haines M A Pericak-Vance G Schellenberg John Sai Keong Kauwe K L Lunetta L A Farrer Justina P. Tavana N. O. Jensen M Rosene Perry Gene Ridge John Sai Keong Kauwe C. M. Karch Corinne E. Sexton Mark Ebbert Ryan H. Miller Meganne Ferrel Jo Ann T. Tschanz Christopher D. Corcoran Alzheimer's Disease Neuroimaging Initiative Perry Gene Ridge John Sai Keong Kauwe Celeste Sassi M A Nalls Perry Gene Ridge R Gibbs M K Lupton C Troakes K Lunnon S Al-Sarraj K S Brown C Medway J Lord J Turton J Bras ARUK Consortium, S Blumenau M Thielke C Josties D Freyer A Dietrich M Hammer M Baier U Dirnagl K Morgan J F Powell John Sai Keong Kauwe C Cruchaga Goate, A M Singleton, A B R Guerreiro A Hodges J Hardy Perry Gene Ridge John Sai Keong Kauwe T J Maxwell C Corcoran J L Del-Aguila J P Budde Y Deming C Cruchaga A M Goate John Sai Keong Kauwe L A Cannon-Albright S Dintelman T Maness J Cerny A Thomas S Backus J M Farnham C C Teerlink J Contreras John Sai Keong Kauwe L J Meyer V Anttila B Bulik-Sullivan H K Finucane R K Walters J Bras L Duncan V Escott-Price G J Falcone P Gormley R Malik N A Patsopoulos S Ripke Z Wei D Yu P H Lee P Turley B Grenier-Boley V Chouraki Y Kamatani C Berr L Letenneur D Hannequin P Amouyel A Boland J F Deleuze E Duron B N Vardarajan C Reitz A M Goate M J Huentelman M I Kamboh E B Larson E Rogaeva P St George-Hyslop H Hakonarson W A Kukull L A Farrer L L Barnes T G Beach F Y Demirci E Head C M Hulette G A Jicha John Sai Keong Kauwe J A Kaye J B Leverenz A I Levey A P Lieberman V S Pankratz W W Poon J F Quinn A J Saykin L S Schneider A G Smith J A Sonnen R A Stern V M Van Deerlin L J Van Eldik D Harold G Russo D C Rubinsztein A Bayer M Tsolaki P Proitsi N C Fox H Hampel M J Owen S Mead P Passmore K Morgan M M Nöthen M Rossor M K Lupton P Hoffmann J Kornhuber B Lawlor A McQuillin A Al-Chalabi J C Bis A Ruiz M Boada S Seshadri A Beiser K Rice S J van der Lee P L De Jager D H Geschwind M Riemenschneider S Riedel-Heller J I Rotter G Ransmayr B T Hyman C Cruchaga M Alegret B Winsvold P Palta K H Farh E Cuenca-Leon N Furlotte T Kurth L Ligthart G M Terwindt T Freilinger C Ran S D Gordon G Borck Adams HHH T Lehtimäki J Wedenoja J E Buring M Schürks M Hrafnsdottir J J Hottenga B Penninx V Artto M Kaunisto S Vepsäläinen N G Martin G W Montgomery M I Kurki E Hämäläinen H Huang J Huang C Sandor C Webber B Muller-Myhsok S Schreiber V Salomaa E Loehrer H Göbel A Macaya P Pozo-Rosich T Hansen T Werge J Kaprio A Metspalu C Kubisch M D Ferrari A C Belin van den Maagdenberg AMJM J A Zwart D Boomsma N Eriksson J Olesen D I Chasman D R Nyholt A Avbersek L Baum S Berkovic J Bradfield R Buono C B Catarino P Cossette P De Jonghe C Depondt D Dlugos T N Ferraro J French H Hjalgrim J Jamnadas-Khoda R Kälviäinen W S Kunz H Lerche C Leu D Lindhout W Lo D Lowenstein M McCormack R S Møller A Molloy P W Ng K Oliver M Privitera R Radtke A K Ruppert T Sander S Schachter C Schankin I Scheffer S Schoch S M Sisodiya P Smith M Sperling P Striano R Surges G N Thomas F Visscher C D Whelan F Zara E L Heinzen A Marson F Becker H Stroink F Zimprich T Gasser R Gibbs P Heutink M Martinez H R Morris M Sharma M Ryten K Y Mok S Pulit S Bevan E Holliday J Attia T Battey G Boncoraglio V Thijs W M Chen B Mitchell P Rothwell P Sharma C Sudlow A Vicente H Markus C Kourkoulis J Pera M Raffeld S Silliman V Boraska Perica L M Thornton L M Huckins N William Rayner C M Lewis M Gratacos F Rybakowski A Keski-Rahkonen A Raevuori J I Hudson T Reichborn-Kjennerud P Monteleone A Karwautz K Mannik J H Baker J K O'Toole S E Trace Davis OSP S G Helder S Ehrlich B Herpertz-Dahlmann U N Danner A A van Elburg M Clementi M Forzan E Docampo J Lissowska J Hauser A Tortorella M Maj F Gonidakis K Tziouvas H Papezova Z Yilmaz G Wagner S Cohen-Woods S Herms A Julià R Rabionet D M Dick S Ripatti O A Andreassen T Espeseth A J Lundervold V M Steen D Pinto S W Scherer H Aschauer A Schosser L Alfredsson L Padyukov K A Halmi J Mitchell M Strober A W Bergen W Kaye J P Szatkiewicz B Cormand J A Ramos-Quiroga C Sánchez-Mora M Ribasés M Casas A Hervas M J Arranz J Haavik T Zayats S Johansson N Williams A Dempfle A Rothenberger J Kuntsi R D Oades T Banaschewski B Franke J K Buitelaar A Arias Vasquez A E Doyle A Reif K P Lesch C Freitag O Rivero H Palmason M Romanos K Langley M Rietschel S H Witt S Dalsgaard A D Børglum I Waldman B Wilmot N Molly Bau CHD J Crosbie R Schachar S K Loo J J McGough E H Grevet S E Medland E Robinson L A Weiss E Bacchelli A Bailey V Bal A Battaglia C Betancur P Bolton R Cantor P Celestino-Soper G Dawson S De Rubeis F Duque A Green S M Klauck M Leboyer P Levitt E Maestrini S Mane D M De-Luca J Parr R Regan A Reichenberg S Sandin J Vorstman T Wassink E Wijsman E Cook S Santangelo R Delorme B Rogé T Magalhaes D Arking T G Schulze R C Thompson J Strohmaier K Matthews I Melle D Morris D Blackwood A McIntosh S E Bergen M Schalling S Jamain A Maaser S B Fischer C S Reinbold J M Fullerton J Guzman-Parra F Mayoral P R Schofield S Cichon T W Mühleisen F Degenhardt J Schumacher M Bauer P B Mitchell E S Gershon J Rice J B Potash P P Zandi N Craddock I N Ferrier M Alda G A Rouleau G Turecki R Ophoff C Pato A Anjorin E Stahl M Leber P M Czerski C Cruceanu I R Jones D Posthuma Andlauer TFM A J Forstner F Streit B T Baune T Air G Sinnamon N R Wray D J MacIntyre D Porteous G Homuth M Rivera J Grove C M Middeldorp I Hickie M Pergadia D Mehta J H Smit R Jansen E de Geus E Dunn Q S Li M Nauck R A Schoevers A T Beekman J A Knowles A Viktorin P Arnold C L Barr G Bedoya-Berrio O J Bienvenu H Brentani C Burton B Camarena C Cappi D Cath M Cavallini D Cusi S Darrow D Denys E M Derks A Dietrich T Fernandez M Figee N Freimer G Gerber M Grados E Greenberg G L Hanna A Hartmann M E Hirschtritt P J Hoekstra A Huang C Huyser C Illmann M Jenike S Kuperman B Leventhal C Lochner G J Lyon F Macciardi M Madruga-Garrido I A Malaty A Maras L McGrath E C Miguel P Mir G Nestadt H Nicolini M S Okun A Pakstis P Paschou J Piacentini C Pittenger K Plessen V Ramensky E M Ramos V Reus M A Richter M A Riddle M M Robertson V Roessner M Rosário J F Samuels P Sandor D J Stein F Tsetsos F Van Nieuwerburgh S Weatherall J R Wendland T Wolanczyk Y Worbe G Zai F S Goes N McLaughlin P S Nestadt H J Grabe C Depienne A Konkashbaev N Lanzagorta A Valencia-Duarte E Bramon N Buccola W Cahn M Cairns S A Chong D Cohen B Crespo-Facorro J Crowley M Davidson L DeLisi T Dinan G Donohoe E Drapeau J Duan L Haan D Hougaard S Karachanak-Yankova A Khrunin J Klovins V Kučinskas J Lee Chee Keong S Limborska C Loughland J Lönnqvist B Maher M Mattheisen C McDonald K C Murphy I Nenadic J van Os C Pantelis M Pato T Petryshen D Quested P Roussos A R Sanders U Schall S G Schwab K Sim H C So E Stögmann M Subramaniam D Toncheva J Waddington J Walters M Weiser W Cheng R Cloninger D Curtis P V Gejman F Henskens M Mattingsdal S Y Oh R Scott B Webb G Breen C Churchhouse C M Bulik M Daly M Dichgans S V Faraone R Guerreiro P Holmans K S Kendler B Koeleman C A Mathews A Price J Scharf P Sklar J Williams N W Wood C Cotsapas A Palotie J W Smoller P Sullivan J Rosand A Corvin B M Neale Perry Gene Ridge Mark E. Wadsworth Justin B. Miller Andrew J. Saykin Robert C. Green the Alzheimer's Disease Neuroimaging Initiative John Sai Keong Kauwe Lynette A. Averill Christopher L. Averill Lyndsay A. Staley J. L. Osawa-Kirk John Sai Keong Kauwe Patricia Henri-Barrus Perry G Ridge Celeste M Karch Simon Hsu Ivan Arano Craig C Teerlink Mark T W Ebbert Jose D Gonzalez Murcia James M Farnham Anna R Damato Mariet Allen Xue Wang Oscar Harari Victoria M Fernandez Rita Guerreiro Jose Bras John Hardy Ronald Munger Maria Norton Celeste Sassi Andrew Singleton Steven G Younkin Dennis W Dickson Todd E Golde Nathan D Price Nilufer Ertekin-Taner Carlos Cruchaga Alison M Goate Christopher Corcoran JoAnn Tschanz Lisa A Cannon-Albright John S K Kauwe Alzheimer's Disease Neuroimaging Initiative R Sims S J van der Lee A C Naj C Bellenguez N Badarinarayan J Jakobsdottir B W Kunkle A Boland R Raybould J C Bis E R Martin B Grenier-Boley S Heilmann-Heimbach V Chouraki A B Kuzma K Sleegers M Vronskaya A Ruiz R R Graham R Olaso P Hoffmann M L Grove B N Vardarajan M Hiltunen M M Nöthen C C White K L Hamilton-Nelson J Epelbaum W Maier S H Choi G W Beecham C Dulary S Herms A V Smith C C Funk C Derbois A J Forstner S Ahmad H Li D Bacq D Harold C L Satizabal O Valladares A Squassina R Thomas J A Brody L Qu P Sánchez-Juan T Morgan F J Wolters Y Zhao F S Garcia N Denning M Fornage J Malamon Naranjo MCD E Majounie T H Mosley B Dombroski D Wallon M K Lupton J Dupuis P Whitehead L Fratiglioni C Medway X Jian S Mukherjee L Keller K Brown H Lin L B Cantwell F Panza B McGuinness S Moreno-Grau J D Burgess V Solfrizzi P Proitsi H H Adams M Allen D Seripa P Pastor L A Cupples N D Price D Hannequin A Frank-García D Levy P Chakrabarty P Caffarra I Giegling A S Beiser V Giedraitis H Hampel M E Garcia X Wang L Lannfelt P Mecocci G Eiriksdottir P K Crane F Pasquier V Boccardi I Henández R C Barber M Scherer L Tarraga P M Adams M Leber Y Chen M S Albert S Riedel-Heller V Emilsson D Beekly A Braae R Schmidt D Blacker C Masullo H Schmidt R S Doody G Spalletta WTL, Jr T J Fairchild P Bossù O L Lopez M P Frosch E Sacchinelli B Ghetti Q Yang R M Huebinger F Jessen S Li M I Kamboh J Morris O Sotolongo-Grau M J Katz C Corcoran M Dunstan A Braddel C Thomas A Meggy R Marshall A Gerrish J Chapman M Aguilar S Taylor M Hill M D Fairén A Hodges B Vellas H Soininen I Kloszewska M Daniilidou J Uphill Y Patel J T Hughes J Lord J Turton A M Hartmann R Cecchetti C Fenoglio M Serpente M Arcaro C Caltagirone M D Orfei A Ciaramella S Pichler M Mayhaus W Gu A Lleó J Fortea R Blesa I S Barber K Brookes C Cupidi R G Maletta D Carrell S Sorbi S Moebus M Urbano A Pilotto J Kornhuber P Bosco S Todd D Craig J Johnston M Gill B Lawlor A Lynch N C Fox J Hardy R L Albin L G Apostolova S E Arnold S Asthana C S Atwood C T Baldwin L L Barnes S Barral T G Beach J T Becker E H Bigio T D Bird B F Boeve J D Bowen A Boxer J R Burke J M Burns J D Buxbaum N J Cairns C Cao C S Carlson C M Carlsson R M Carney M M Carrasquillo S L Carroll C C Diaz H C Chui D G Clark D H Cribbs E A Crocco C DeCarli M Dick R Duara D A Evans K M Faber K B Fallon D W Fardo M R Farlow S Ferris T M Foroud D R Galasko M Gearing D H Geschwind J R Gilbert N R Graff-Radford R C Green J H Growdon R L Hamilton L E Harrell L S Honig M J Huentelman C M Hulette B T Hyman G P Jarvik E Abner L W Jin G Jun A Karydas J A Kaye R Kim N W Kowall J H Kramer F M LaFerla J J Lah J B Leverenz A I Levey G Li A P Lieberman K L Lunetta C G Lyketsos D C Marson F Martiniuk D C Mash E Masliah W C McCormick S M McCurry A N McDavid A C McKee M Mesulam B L Miller C A Miller J W Miller J C Morris J R Murrell A J Myers S O'Bryant J M Olichney V S Pankratz J E Parisi H L Paulson W Perry E Peskind A Pierce W W Poon H Potter J F Quinn A Raj M Raskind B Reisberg C Reitz J M Ringman E D Roberson E Rogaeva H J Rosen R N Rosenberg M A Sager A J Saykin J A Schneider L S Schneider W W Seeley A G Smith J A Sonnen S Spina R A Stern R H Swerdlow R E Tanzi T A Thornton-Wells J Q Trojanowski J C Troncoso V M Van Deerlin L J Van Eldik H V Vinters J P Vonsattel S Weintraub K A Welsh-Bohmer K C Wilhelmsen J Williamson T S Wingo R L Woltjer C B Wright C E Yu L Yu F Garzia F Golamaully G Septier S Engelborghs R Vandenberghe P P De Deyn C M Fernadez Y A Benito H Thonberg C Forsell L Lilius A Kinhult-Stählbom L Kilander R Brundin L Concari S Helisalmi A M Koivisto A Haapasalo V Dermecourt N Fievet O Hanon C Dufouil A Brice K Ritchie B Dubois J J Himali C D Keene J Tschanz A L Fitzpatrick W A Kukull M Norton T Aspelund E B Larson R Munger J I Rotter R B Lipton M J Bullido A Hofman T J Montine E Coto E Boerwinkle R C Petersen V Alvarez F Rivadeneira E M Reiman M Gallo C J O'Donnell J S Reisch A C Bruni D R Royall M Dichgans M Sano D Galimberti P St George-Hyslop E Scarpini D W Tsuang M Mancuso U Bonuccelli A R Winslow A Daniele C K Wu O Peters B Nacmias M Riemenschneider R Heun C Brayne D C Rubinsztein J Bras R Guerreiro A Al-Chalabi C E Shaw J Collinge D Mann M Tsolaki J Clarimón R Sussams S Lovestone M C O'Donovan M J Owen T W Behrens S Mead A M Goate A G Uitterlinden C Holmes C Cruchaga M Ingelsson D A Bennett J Powell T E Golde C Graff P L De Jager K Morgan N Ertekin-Taner O Combarros B M Psaty P Passmore S G Younkin C Berr V Gudnason D Rujescu D W Dickson J F Dartigues A L DeStefano S Ortega-Cubero H Hakonarson D Campion M Boada J K Kauwe L A Farrer C Van Broeckhoven M A Ikram L Jones J L Haines C Tzourio L J Launer V Escott-Price R Mayeux J F Deleuze N Amin P A Holmans M A Pericak-Vance P Amouyel C M van Duijn A Ramirez L S Wang J C Lambert S Seshadri J Williams G D Schellenberg K L Huang E Marcora A A Pimenova A F Di Narzo M Kapoor S C Jin O Harari S Bertelsen B P Fairfax J Czajkowski V Chouraki B Grenier-Boley C Bellenguez Y Deming A McKenzie T Raj A E Renton J Budde A Smith A Fitzpatrick J C Bis A DeStefano Adams HHH M A Ikram S van der Lee J L Del-Aguila M V Fernandez L Ibañez R Sims V Escott-Price R Mayeux J L Haines L A Farrer M A Pericak-Vance J C Lambert C van Duijn L Launer S Seshadri J Williams P Amouyel G D Schellenberg B Zhang I Borecki John Sai Keong Kauwe C Cruchaga K Hao A M Goate Swati Anand Justin M Barnes Sydney A Young Diana M Garcia Harold Dennis Tolley John Sai Keong Kauwe Steven William Graves J Matyi J T Tschanz G B Rattinger C Sanders E K Vernon C Corcoran John Sai Keong Kauwe M Buhusi T Parks M M Mirabel J Kado K Auckland J Nowak A Rautanen A J Mentzer E Marijon X Jouven M L Perman T Cua John Sai Keong Kauwe J B Allen H Taylor K J Robson C M Deane A C Steer Hill AVS Lori B. Allen Fauatea Henry Taylor Adonica I. Kauwe Tawny Larsen Ariel A. Hippen Marvin Allen John Sai Keong Kauwe Y Deming Z Li M Kapoor O Harari J L Del-Aguila K Black D Carrell Y Cai M V Fernandez J Budde S Ma B Saef B Howells K L Huang S Bertelsen A M Fagan D M Holtzman J C Morris S Kim A J Saykin P L De Jager M Albert A Moghekar R O'Brien M Riemenschneider R C Petersen K Blennow H Zetterberg L Minthon V M Van Deerlin V M Lee L M Shaw J Q Trojanowski G Schellenberg J L Haines R Mayeux M A Pericak-Vance L A Farrer E R Peskind G Li A F Di Narzo John Sai Keong Kauwe A M Goate C Cruchaga D W Fardo Y Katsumata John Sai Keong Kauwe Y Deming O Harari C Cruchaga P T Nelson N Z Steele J S Carr L W Bonham E G Geier V Damotte Z A Miller R S Desikan K L Boehme S Mukherjee P K Crane John Sai Keong Kauwe J H Kramer B L Miller G Coppola J A Hollenbach Y Huang J S Yokoyama S Mukherjee J C Russell D T Carr J D Burgess M Allen D J Serie K L Boehme John Sai Keong Kauwe A C Naj D W Fardo D W Dickson T J Montine N Ertekin-Taner M R Kaeberlein P K Crane G R Jun J Chung J Mez R Barber G W Beecham D A Bennett J D Buxbaum G S Byrd M M Carrasquillo P K Crane C Cruchaga P De Jager N Ertekin-Taner D Evans M D Fallin T M Foroud R P Friedland A M Goate N R Graff-Radford H Hendrie K S Hall K L Hamilton-Nelson R Inzelberg M I Kamboh John Sai Keong Kauwe W A Kukull B W Kunkle R Kuwano E B Larson M W Logue J J Manly E R Martin T J Montine S Mukherjee A Naj E M Reiman C Reitz R Sherva P H St George-Hyslop T Thornton S G Younkin B N Vardarajan L S Wang J R Wendlund A R Winslow J Haines R Mayeux M A Pericak-Vance G Schellenberg K L Lunetta L A Farrer P C Haddick J L Larson N Rathore T R Bhangale Q T Phung K Srinivasan D V Hansen J R Lill M A Pericak-Vance J Haines L A Farrer John Sai Keong Kauwe G D Schellenberg C Cruchaga A M Goate T W Behrens R J Watts R R Graham J S Kaminker M van der Brug G. Jun C. A. Ibrahim-Verbaas M. Vronskaya J C Lambert J. Chung A. C. Naj B. W. Kunkle L. S. Wang J. C. Bis C. Bellenguez D. Harold K. L. Lunetta A. L. Destefano B. Grenier-Boley R. Sims G. W. Beecham A. V. Smith V. Chouraki K. L. Hamilton-Nelson M. A. Ikram N. Fievet N. Denning E. R. Martin H. Schmidt Y. Kamatani M. L. Dunstan O. Valladares A. R. Laza D. Zelenika A. Ramirez T. M. Foroud S. H. Choi A. Boland T. Becker W. A. Kukull S. J. van der Lee F. Pasquier C. Cruchaga D. Beekly A. L. Fitzpatrick O. Hanon M. Gill R. Barber V. Gudnason D. Campion S. Love D. A. Bennett N. Amin C. Berr M. Tsolaki J. D. Buxbaum O. L. Lopez V. Deramecourt N. C. Fox L. B. Cantwell L. Tarraga C. Dufouil J. Hardy P. K. Crane G. Eiriksdottir D. Hannequin R. Clarke D. Evans T. H. Mosley L. Letenneur C. Brayne W. Maier P. De Jager V. Emilsson J. F. Dartigues H. Hampel M. I. Kamboh R. F. de Bruijn C. Tzourio P. Pastor E. B. Larson J. I. Rotter M. C. O'Donovan T. J. Montine M. A. Nalls S. Mead E. M. Reiman P. V. Jonsson C. Holmes P. H. St George-Hyslop M. Boada P. Passmore Schmidt R. Wendland K. Morgan A. R. Winslow J. F. Powell M. Carasquillo S. G. Younkin J. Jakobsdottir John Sai Keong Kauwe K. C. Wilhelmsen D. Rujescu M. M. Nothen A. Hofman L. Jones Consortium I J. L. Haines B. M. Psaty C. Van Broeckhoven P. Holmans L. J. Launer R. Mayeux M. Lathrop A. M. Goate V. Escott-Price S. Seshadri M. A. Pericak-Vance P. Amouyel J. Williams C. M. van Duijn G. D. Schellenberg L. A. Farrer Y. Deming J. Xia Y. Cai J. Lord P. Holmans S. Bertelsen D. Holtzman J. C. Morris K. Bales E. H. Pickering A. Goate John Sai Keong Kauwe C. Cruchaga Celeste Sassi Michael A Nalls Perry Gene Ridge R Gibbs Jinhui Ding Michelle K Lupton Claire Troakes Katie Lunnon Safa Al-Sarraj Kristelle S Brown John Sai Keong Kauwe others Perry Gene Ridge Kaitlyn B Hoyt Kevin Boehme Shubhabrata Mukherjee Paul K Crane Jonathan L Haines Richard Mayeux Lindsay A Farrer Margaret A Pericak-Vance Gerard D Schellenberg John Sai Keong Kauwe others G. I. Allen N. Amoroso C. Anghel V. Balagurusamy C. J. Bare D. Beaton R. Bellotti D. A. Bennett K. Boehme P. C. Boutros L. Caberlotto C. Caloian F. Campbell E. C. Neto Y. C. Chang B. Chen C. Y. Chen T. Y. Chien T. Clark S. Das C. Davatzikos J. Deng D. Dillenberger R. J. Dobson Q. Dong J. Doshi D. Duma R. Errico G. Erus E. Everett D. W. Fardo S. H. Friend H. Frohlich J. Gan P. St George-Hyslop S. S. Ghosh E. Glaab R. C. Green Y. Guan M. Y. Hong C. Huang J. Hwang J. Ibrahim P. Inglese Q. Jiang Y. Katsumata John Sai Keong Kauwe A. Klein D. Kong R. Krause E. Lalonde M. Lauria E. Lee X. Lin Z. Liu J. Livingstone B. A. Logsdon S. Lovestone A. Lyappan M. Ma A. Malhotra L. M. Mangravite T. J. Maxwell E. Merrill J. Nagorski A. Namasivayam M. Narayan M. Naz S. J. Newhouse T. C. Norman R. N. Nurtdinov Y. J. Oyang Y. Pawitan S. Peng M. A. Peters Stephen R Piccolo P. Praveen C. Priami V. Y. Sabelnykova P. Senger X. Shen A. Simmons A. Sotiras G. Stolovitzky S. Tangaro A. Tateo Y. A. Tung N. J. Tustison E. Varol G. Vradenburg M. W. Weiner G. Xiao L. Xie Y. Xie J. Xu H. Yang X. Zhan Y. Zhou F. Zhu H. Zhu S. Zhu Dipti Shah Frederick Rohlfing Swati Anand W Evan Johnson MeiHwa TB Alvarez Jesse Cobell Jackson King Sydney A Young John Sai Keong Kauwe Steven William Graves Mark TW Ebbert Mark E Wadsworth Lyndsay A Staley Kaitlyn L Hoyt Brandon Pickett Justin Miller John Duce John Sai Keong Kauwe Perry Gene Ridge John Sai Keong Kauwe A. M. Goate Yuetiva Deming Jian Xia Yefei Cai Jenny Lord Jorge L Del-Aguila Maria Victoria Fernandez David Carrell Kathleen Black John Budde ShengMei Ma Perry Gene Ridge John Sai Keong Kauwe others Lyndsay A Staley Mark TW Ebbert Sheradyn Parker Matthew Bailey Perry Gene Ridge Alison M Goate John Sai Keong Kauwe J. Chapuis A. Flaig B. Grenier-Boley F. Eysert V. Pottiez G. Deloison A. Vandeputte A. M. Ayral T. Mendes S. Desai A. M. Goate John Sai Keong Kauwe F. Leroux A. Herledan F. Demiautte C. Bauer F. Checler R. C. Petersen K. Blennow H. Zetterberg L. Minthon V. M. Van Deerlin V. M. Lee L. M. Shaw J. Q. Trojanowski M. Albert A. Moghekar R. O'Brien E. R. Peskind N. Malmanche G. D. Schellenberg P. Dourlen O. R. Song C. Cruchaga P. Amouyel B. Deprez P. Brodin J. C. Lambert C. Sassi P. G. Ridge M. A. Nalls R. Gibbs J. Ding M. K. Lupton C. Troakes K. Lunnon S. Al-Sarraj K. S. Brown C. Medway J. Lord J. Thurton K. Morgan J. F. Powell John Sai Keong Kauwe C. Cruchaga J. Bras A. M. Goate A. B. Singleton R. Guerreiro J. Hardy Mark TW Ebbert Kevin L Boehme Mark E Wadsworth Lyndsay A Staley for the Alzheimer’s Disease Neuroimaging Initiative Alzheimer’s Disease Genetics Consortium Shubhabrata Mukherjee Paul K Crane Perry G Ridge John SK Kauwe A. A. Hippen M. T. W. Ebbert M. C. Norton J. T. Tschanz R. G. Munger C. D. Corcoran John Sai Keong Kauwe E. Oldoni G. G. Fumagalli M. Serpente C. Fenoglio M. Scarioni A. Arighi G. Bruno G. Talarico A. Confaloni P. Piscopo B. Nacmias S. Sorbi I. Rainero E. Rubino L. Pinessi G. Binetti R. Ghidoni L. Benussi G. Grande B. Arosio D. Bursey John Sai Keong Kauwe S. M. Cioffi M. Arcaro D. Mari C. Mariani E. Scarpini D. Galimberti J. Mez S. Mukherjee T. Thornton D. W. Fardo E. Trittschuh S. Sutti R. Sherva John Sai Keong Kauwe A. C. Naj G. W. Beecham A. Gross A. J. Saykin R. C. Green P. K. Crane Lyndsay A Staley Mark TW Ebbert Daniel Bunker Matthew Bailey Perry Gene Ridge Alison M Goate John Sai Keong Kauwe Mark TW Ebbert Lyndsay A Staley Joshua Parker Sheradyn Parker Matthew Bailey Perry Gene Ridge Alison M Goate John Sai Keong Kauwe Cindee E Perry Shawn D Gale Lance David Erickson Eric Wilson Brent Nielsen John Sai Keong Kauwe Dawson Ward Hedges Ting Chen Timothy M Moore Mark T Ebbert Natalie L McVey Steven R Madsen David M Hallowell Alex M Harris Ryan P Mackay Robin E Char Chad R Hancock Jason Merrell Hansen John Sai Keong Kauwe David Morley Thomson S. D. Ostergaard S. Mukherjee S. J. Sharp P. Proitsi L. A. Lotta F. Day Perry KL Boehme S. Walter John Sai Keong Kauwe L. E. Gibbons C. Alzheimer's Disease Genetics Consortium G Consortium EP-I E. B. Larson J. F. Powell C. Langenberg P. K. Crane N. J. Wareham R. A. Scott A. J. Saykin L. Shen X. Yao S. Kim K. Nho S. L. Risacher V. K. Ramanan T. M. Foroud K. M. Faber N. Sarwar L. M. Munsie X. Hu H. D. Soares S. G. Potkin P. M. Thompson John Sai Keong Kauwe R. Kaddurah-Daouk R. C. Green A. W. Toga M. W. Weiner I. Alzheimer's Disease Neuroimaging S. Mukherjee S. Walter John Sai Keong Kauwe A. J. Saykin D. A. Bennett P. K. Crane M. M. Glymour L. S. Wang A. C. Naj R. R. Graham P. K. Crane B. W. Kunkle C. Cruchaga J. D. Murcia L. Cannon-Albright C. T. Baldwin H. Zetterberg K. Blennow W. A. Kukull K. M. Faber N. Schupf M. C. Norton J. T. Tschanz R. G. Munger C. D. Corcoran E. Rogaeva C. Alzheimer's Disease Genetics C. F. Lin B. A. Dombroski L. B. Cantwell A. Partch O. Valladares H. Hakonarson P. St George-Hyslop R. C. Green A. M. Goate T. M. Foroud R. M. Carney E. B. Larson T. W. Behrens John Sai Keong Kauwe J. L. Haines L. A. Farrer M. A. Pericak-Vance R. Mayeux G. D. Schellenberg National Institute on Aging-Late-Onset Alzheimer's Disease Family S M. S. Albert R. L. Albin L. G. Apostolova S. E. Arnold R. Barber M. Barmada L. L. Barnes T. G. Beach J. T. Becker G. W. Beecham D. Beekly D. A. Bennett E. H. Bigio T. D. Bird D. Blacker B. F. Boeve J. D. Bowen A. Boxer Buxbaum JD Burke N. J. Cairns C. Cao C. S. Carlson S. L. Carroll H. C. Chui D. G. Clark D. H. Cribbs E. A. Crocco C. DeCarli S. T. DeKosky F. Y. Demirci M. Dick D. W. Dickson R. Duara N. Ertekin-Taner K. B. Fallon M. R. Farlow S. Ferris M. P. Frosch D. R. Galasko M. Ganguli M. Gearing D. H. Geschwind B. Ghetti Glass JD Gilbert N. R. Graff-Radford J. H. Growdon R. L. Hamilton K. L. Hamilton-Nelson L. E. Harrell E. Head L. S. Honig C. M. Hulette B. T. Hyman G. P. Jarvik G. A. Jicha L. W. Jin G. Jun G. Jun M. I. Kamboh A. Karydas J. A. Kaye R. Kim E. H. Koo N. W. Kowall J. H. Kramer F. M. LaFerla J. J. Lah J. B. Leverenz A. I. Levey G. Li A. P. Lieberman O. L. Lopez K. L. Lunetta C. G. Lyketsos W. J. Mack D. C. Marson E. R. Martin F. Martiniuk D. C. Mash E. Masliah W. C. McCormick S. M. McCurry A. N. McDavid A. C. McKee W. M. Mesulam B. L. Miller C. A. Miller J. W. Miller T. J. Montine J. C. Morris Olichney JM Murrell J. E. Parisi W. Perry E. Peskind R. C. Petersen A. Pierce W. W. Poon H. Potter J. F. Quinn A. Raj M. Raskind E. M. Reiman B. Reisberg C. Reitz J. M. Ringman E. D. Roberson H. J. Rosen R. N. Rosenberg M. Sano A. J. Saykin J. A. Schneider L. S. Schneider W. W. Seeley A. G. Smith J. A. Sonnen S. Spina R. A. Stern R. E. Tanzi T. A. Thornton-Wells J. Q. Trojanowski J. C. Troncoso D. W. Tsuang V. M. Van Deerlin L. J. Van Eldik B. N. Vardarajan H. V. Vinters J. P. Vonsattel S. Weintraub K. A. Welsh-Bohmer J. Williamson S. Wishnek R. L. Woltjer C. B. Wright S. G. Younkin C. E. Yu L. Yu C Lythgoe A Perkes M Peterson C Schmutz M Leary M T Ebbert Perry Gene Ridge M C Norton J T Tschanz R G Munger C D Corcoran John Sai Keong Kauwe Mark TW Ebbert Perry G Ridge John SK Kauwe Matthew Seeley Mark Joel Clement Christophe Gerard Giraud-Carrier Quinn O Snell Paul Bodily Stanley Fujimoto John Sai Keong Kauwe Perry Gene Ridge Dawson Ward Hedges Cynthia Perry Brent Nielsen Brian Jeffs Malvina Salash Ronald L Hager Lance David Erickson Vaughn R A Call Pat Roy Vehrs John Sai Keong Kauwe L. Shen P. M. Thompson S. G. Potkin L. Bertram L. A. Farrer T. M. Foroud R. C. Green X. Hu M. J. Huentelman S. Kim John Sai Keong Kauwe Q. Li E. Liu F. Macciardi J. H. Moore L. Munsie K. Nho V. K. Ramanan S. L. Risacher D. J. Stone S. Swaminathan A. W. Toga M. W. Weiner A. J. Saykin for the Alzheimer’s Disease Neuroimaging Initiative. P. Dayton Merrill Kauwe John Sai Keong Kauwe M. Feilmeier J. Hirschi M. T. Ebbert Perry Gene Ridge A. R. Wilson A. R. Sharp M. Bailey M. C. Norton J. T. Tschanz R. G. Munger C. D. Corcoran John Sai Keong Kauwe D. Peterson C. Munger J. Crowley C. Corcoran C. Cruchaga A. M. Goate M. C. Norton R. C. Green Alzheimer's Disease Neuroimaging Initiative R. G. Munger J. C. Breitner K. A. Welsh-Bohmer C. Lyketsos J. Tschanz John Sai Keong Kauwe A C Naj G Jun C Reitz B W Kunkle W Perry Y S Park G W Beecham R A Rajbhandary K L Hamilton-Nelson L S Wang John Sai Keong Kauwe M J Huentelman A J Myers T D Bird B F Boeve C T Baldwin G P Jarvik P K Crane E Rogaeva M M Barmada F Y Demirci C Cruchaga P L Kramer N Ertekin-Taner J Hardy N R Graff-Radford R C Green E B Larson P H St George-Hyslop J D Buxbaum D A Evans J A Schneider K L Lunetta M I Kamboh A J Saykin E M Reiman P L De Jager D A Bennett J C Morris T J Montine A M Goate D Blacker D W Tsuang H Hakonarson W A Kukull T M Foroud E R Martin J L Haines R P Mayeux L A Farrer G D Schellenberg M A Pericak-Vance M S Albert R L Albin L G Apostolova S E Arnold R Barber L L Barnes T G Beach J T Becker D Beekly E H Bigio J D Bowen A Boxer J R Burke N J Cairns L B Cantwell C Cao C S Carlson R M Carney M M Carrasquillo S L Carroll H C Chui D G Clark J Corneveaux D H Cribbs E A Crocco C DeCarli S T DeKosky M Dick D W Dickson R Duara K M Faber K B Fallon M R Farlow S Ferris M P Frosch D R Galasko M Ganguli M Gearing D H Geschwind B Ghetti J R Gilbert J D Glass J H Growdon R L Hamilton L E Harrell E Head L S Honig C M Hulette B T Hyman G A Jicha L W Jin A Karydas J A Kaye R Kim E H Koo N W Kowall J H Kramer F M LaFerla J J Lah J B Leverenz A I Levey G Li A P Lieberman Chiao Fen Lin O L Lopez C G Lyketsos W J Mack F Martiniuk D C Mash E Masliah W C McCormick S M McCurry A N McDavid A C McKee M Mesulam B L Miller C A Miller J W Miller J R Murrell J M Olichney V S Pankratz J E Parisi H L Paulson E Peskind R C Petersen A Pierce W W Poon H Potter J F Quinn A Raj M Raskind B Reisberg J M Ringman E D Roberson H J Rosen R N Rosenberg M Sano Lon S Schneider W W Seeley A G Smith J A Sonnen S Spina R A Stern R E Tanzi T A Thornton-Wells J Q Trojanowski J C Troncoso O Valladares V M Van Deerlin L J Van Eldik B N Vardarajan H V Vinters J P Vonsattel S Weintraub K A Welsh-Bohmer J Williamson S Wishnek R L Woltjer C B Wright S G Younkin C E Yu L Yu John Sai Keong Kauwe M H Bailey Perry Gene Ridge R Perry M E Wadsworth K L Hoyt L A Staley C M Karch O Harari C Cruchaga B J Ainscough K Bales E H Pickering S Bertelsen A M Fagan D M Holtzman J C Morris A M Goate M Allen M Kachadoorian Z Quicksall F Zou H S Chai C Younkin J E Crook V S Pankratz M M Carrasquillo S Krishnan T Nguyen L Ma K Malphrus S Lincoln G Bisceglio C P Kolbert J Jen S Mukherjee John Sai Keong Kauwe P K Crane J L Haines R Mayeux M A Pericak-Vance L A Farrer G D Schellenberg J E Parisi R C Petersen N R Graff-Radford D W Dickson S G Younkin N Ertekin-Taner V Escott-Price C Bellenguez L S Wang S H Choi D Harold L Jones P Holmans A Gerrish A Vedernikov A Richards A L DeStefano J C Lambert C A Ibrahim-Verbaas A C Naj R Sims G Jun J C Bis G W Beecham B Grenier-Boley G Russo T A Thornton-Wells N Denning A V Smith V Chouraki C Thomas M A Ikram D Zelenika B N Vardarajan Y Kamatani C F Lin H Schmidt B Kunkle M L Dunstan M Vronskaya A D Johnson A Ruiz M T Bihoreau C Reitz F Pasquier P Hollingworth O Hanon A L Fitzpatrick J D Buxbaum D Campion P K Crane C Baldwin T Becker V Gudnason C Cruchaga D Craig N Amin C Berr O L Lopez P L De Jager V Deramecourt J A Johnston D Evans S Lovestone L Letenneur I Hernández D C Rubinsztein G Eiriksdottir K Sleegers A M Goate N Fiévet M J Huentelman M Gill K Brown M I Kamboh L Keller P Barberger-Gateau B McGuinness E B Larson A J Myers C Dufouil S Todd D Wallon S Love E Rogaeva J Gallacher P S George-Hyslop J Clarimon A Lleo A Bayer D W Tsuang L Yu M Tsolaki P Bossù G Spalletta P Proitsi J Collinge S Sorbi F S Garcia N C Fox J Hardy M C Naranjo P Bosco R Clarke C Brayne D Galimberti E Scarpini U Bonuccelli M Mancuso G Siciliano S Moebus P Mecocci M D Zompo W Maier H Hampel A Pilotto A Frank-García F Panza V Solfrizzi P Caffarra B Nacmias W Perry M Mayhaus L Lannfelt H Hakonarson S Pichler M M Carrasquillo M Ingelsson D Beekly V Alvarez F Zou O Valladares S G Younkin E Coto K L Hamilton-Nelson W Gu C Razquin P Pastor I Mateo M J Owen K M Faber P V Jonsson O Combarros M C O'Donovan L B Cantwell H Soininen D Blacker S Mead T H Mosley, Jr D A Bennett T B Harris L Fratiglioni C Holmes R F de Bruijn P Passmore T J Montine K Bettens J I Rotter A Brice K Morgan T M Foroud W A Kukull D Hannequin J F Powell M A Nalls K Ritchie K L Lunetta John Sai Keong Kauwe E Boerwinkle M Riemenschneider M Boada M Hiltunen E R Martin R Schmidt D Rujescu J F Dartigues R Mayeux C Tzourio A Hofman M M Nöthen C Graff B M Psaty J L Haines M Lathrop M A Pericak-Vance L J Launer C Van Broeckhoven L A Farrer C M van Duijn A Ramirez S Seshadri G D Schellenberg P Amouyel J Williams B A Benitez S C Jin R Guerreiro R Graham J Lord D Harold R Sims J C Lambert J R Gibbs J Bras C Sassi O Harari S Bertelsen M K Lupton J Powell C Bellenguez K Brown C Medway P C Haddick M P van der Brug T Bhangale W Ortmann T Behrens R Mayeux M A Pericak-Vance L A Farrer G D Schellenberg J L Haines J Turton A Braae I Barber A M Fagan D M Holtzman J C Morris J Williams John Sai Keong Kauwe P Amouyel K Morgan A Singleton J Hardy A M Goate C Cruchaga C. Cruchaga M. T. Ebbert John Sai Keong Kauwe P. Dayton M. Feilmeier J. Hirschi M. Kauwe John Sai Keong Kauwe Y. Cao P. Wei M. Bailey John Sai Keong Kauwe T. J. Maxwell C Cruchaga C M Karch S C Jin B A Benitez Y Cai R Guerreiro O Harari J Norton J Budde S Bertelsen A T Jeng B Cooper T Skorupa D Carrell D Levitch S Hsu J Choi M Ryten J Hardy M Ryten D Trabzuni M E Weale A Ramasamy C Smith C Sassi J Bras J R Gibbs D G Hernandez M K Lupton J Powell P Forabosco Perry Gene Ridge C D Corcoran J T Tschanz M C Norton R G Munger C Schmutz M Leary F Y Demirci M N Bamne X Wang O L Lopez M Ganguli C Medway J Turton J Lord A Braae I Barber K Brown P Passmore D Craig J Johnston B McGuinness S Todd R Heun H Kölsch P G Kehoe N M Hooper E R Vardy D M Mann S Pickering-Brown K Brown N Kalsheker J Lowe K Morgan A David Smith G Wilcock D Warden C Holmes P Pastor O Lorenzo-Betancor Z Brkanac E Scott E Topol K Morgan E Rogaeva A B Singleton J Hardy M I Kamboh P St George-Hyslop N Cairns J C Morris John Sai Keong Kauwe A M Goate O. Harari C. Cruchaga John Sai Keong Kauwe B. J. Ainscough K. Bales E. H. Pickering S. Bertelsen A. M. Fagan D. M. Holtzman J. C. Morris A. M. Goate A L Gross R Sherva S Mukherjee S Newhouse John Sai Keong Kauwe L M Munsie L B Waterston D A Bennett R N Jones R C Green P K Crane Perry Gene Ridge T J Maxwell S J Foutz M H Bailey C D Corcoran J T Tschanz M C Norton R G Munger E O'Brien R A Kerber R M Cawthon John Sai Keong Kauwe A R Sharp Perry Gene Ridge M H Bailey K L Boehme M C Norton J T Tschanz R G Munger C D Corcoran John Sai Keong Kauwe M T Ebbert M E Wadsworth K L Boehme K L Hoyt A R Sharp B D O'Fallon John Sai Keong Kauwe Perry Gene Ridge J. D. Gonzalez Murcia C. Schmutz C. Munger A. Perkes A. Gustin M. Peterson M. T. Ebbert M. C. Norton J. T. Tschanz R. G. Munger C. D. Corcoran John Sai Keong Kauwe J C Lambert C A Ibrahim-Verbaas D Harold A C Naj R Sims C Bellenguez A L DeStafano J C Bis G W Beecham B Grenier-Boley G Russo T A Thorton-Wells N Jones A V Smith V Chouraki C Thomas M A Ikram D Zelenika B N Vardarajan Y Kamatani Chiao Fent Lin A Gerrish H Schmidt B Kunkle M L Dunstan A Ruiz M T Bihoreau S H Choi C Reitz F Pasquier C Cruchaga D Craig N Amin C Berr O L Lopez P L De Jager V Deramecourt James A Johnston D Evans S Lovestone L Letenneur F J Morón D C Rubinsztein G Eiriksdottir K Sleegers A M Goate N Fiévet M W Huentelman M Gill K Brown M I Kamboh L Keller P Barberger-Gateau B McGuiness E B Larson R Green A J Myers C Dufouil S Todd D Wallon S Love E Rogaeva J Gallacher P St George-Hyslop J Clarimon A Lleo A Bayer D W Tsuang L Yu M Tsolaki P Bossù G Spalletta P Proitsi J Collinge S Sorbi F Sanchez-Garcia N C Fox J Hardy M C Deniz Naranjo P Bosco R Clarke C Brayne D Galimberti M Mancuso F Matthews S Moebus P Mecocci M Del Zompo W Maier H Hampel A Pilotto M Bullido F Panza P Caffarra B Nacmias J R Gilbert M Mayhaus L Lannefelt H Hakonarson S Pichler M M Carrasquillo M Ingelsson D Beekly V Alvarez F Zou O Valladares S G Younkin E Coto K L Hamilton-Nelson W Gu C Razquin P Pastor I Mateo M J Owen K M Faber P V Jonsson O Combarros M C O'Donovan L B Cantwell H Soininen D Blacker S Mead T H Mosley, Jr D A Bennett T B Harris L Fratiglioni C Holmes R F de Bruijn P Passmore T J Montine K Bettens J I Rotter A Brice K Morgan T M Foroud W A Kukull D Hannequin J F Powell M A Nalls K Ritchie K L Lunetta John Sai Keong Kauwe E Boerwinkle M Riemenschneider M Boada M Hiltuenen E R Martin R Schmidt D Rujescu L S Wang J F Dartigues R Mayeux C Tzourio A Hofman M M Nöthen C Graff B M Psaty L Jones J L Haines P A Holmans M Lathrop M A Pericak-Vance L J Launer L A Farrer C M van Duijn C Van Broeckhoven V Moskvina S Seshadri J Williams G D Schellenberg P Amouyel Perry Gene Ridge Shubhabrata Mukherjee Paul K Crane John Sai Keong Kauwe ADGC John Sai Keong Kauwe Perry Gene Ridge N. L. Foster L. A. Cannon-Albright Perry Gene Ridge A. Koop T. J. Maxwell M. H. Bailey R. H. Swerdlow John Sai Keong Kauwe R. A. Honea Perry Gene Ridge M. T. Ebbert John Sai Keong Kauwe C. Cruchaga John Sai Keong Kauwe O. Harari S. C. Jin Y. Cai C. M. Karch B. A. Benitez A. T. Jeng T. Skorupa D. Carrell S. Bertelsen M. Bailey D. McKean J. M. Shulman P. L. De Jager L. Chibnik D. A. Bennett S. E. Arnold D. Harold R. Sims A. Gerrish J. Williams V. M. Van Deerlin V. M. Lee L. M. Shaw J. Q. Trojanowski J. L. Haines R. Mayeux M. A. Pericak-Vance L. A. Farrer G. D. Schellenberg E. R. Peskind D. Galasko A. M. Fagan D. M. Holtzman J. C. Morris GERAD Consortium Alzheimer's Disease Neuroimaging Initiative (ADNI) Alzheimer Disease Genetic Consortium (ADGC) A. M. Goate Derek D. Houston David B. Elzinga Peter J Maughan Scott M. Smith John Sai Keong Kauwe Richard Paul Evans Ryan B. Stinger Dennis K Shiozawa R. Guerreiro A. Wojtas J. Bras M. Carrasquillo E. Rogaeva E. Majounie C. Cruchaga C. Sassi John Sai Keong Kauwe S. Younkin L. Hazrati J. Collinge J. Pocock T. Lashley J. Williams J. C. Lambert P. Amouyel A. Goate R. RAdemakers K. Morgan J. Powell P. St George-Hhyslop A. Singleton J. Hardy the Alzheimer Genetic Analysis Group. Perry G Ridge PG Ridge TJ Maxwell CD Corcoran MC Norton JT Tschanz E. O'Brien RA Kerber RM Cawthon RG Munger John Sai Keong Kauwe C. Cruchaga John Sai Keong Kauwe P. Nowotny K. Bales EH Pickering K. Mayo S. Bertelsen A. Hinrichs Alzheimer's Disease Neuroimaging Initiative AM Fagan DM Holtzman JC Morris AM Goate A Gerrish G Russo A Richards V Moskvina D Ivanov D Harold R Sims R Abraham P Hollingworth J Chapman M Hamshere J S Pahwa K Dowzell A Williams N Jones C Thomas A Stretton A R Morgan S Lovestone J Powell P Proitsi M K Lupton C Brayne D C Rubinsztein M Gill B Lawlor A Lynch K Morgan K S Brown P A Passmore D Craig B McGuinness S Todd J A Johnston C Holmes D Mann A D Smith S Love P G Kehoe J Hardy S Mead N Fox M Rossor J Collinge W Maier F Jessen H Kölsch R Heun B Schürmann H van den Bussche I Heuser J Kornhuber J Wiltfang M Dichgans L Frölich H Hampel M Hüll D Rujescu A M Goate John Sai Keong Kauwe C Cruchaga P Nowotny J C Morris K Mayo G Livingston N J Bass H Gurling A McQuillin R Gwilliam P Deloukas G Davies S E Harris J M Starr I J Deary A Al-Chalabi C E Shaw M Tsolaki A B Singleton R Guerreiro T W Mühleisen M M Nöthen S Moebus K H Jöckel N Klopp H E Wichmann M M Carrasquillo V S Pankratz S G Younkin L Jones P A Holmans M C O'Donovan M J Owen J Williams E. T. Overton John Sai Keong Kauwe R. Paul K. Tashima D. F. Tate P. Patel C. C. Carpenter D. Patty J. T. Brooks D. B. Clifford Carlos Cruchaga Petra Nowotny John Sai Keong Kauwe Perry G. Ridge Kevin Mayo Sarah Bertelsen Anthony Hinrichs Anne M. Fagan David M. Holtzman John C. Morris Alison M. Goate P Hollingworth D Harold R Sims A Gerrish J C Lambert M M Carrasquillo R Abraham M L Hamshere J S Pahwa V Moskvina K Dowzell N Jones A Stretton C Thomas A Richards D Ivanov C Widdowson J Chapman S Lovestone J Powell P Proitsi M K Lupton C Brayne D C Rubinsztein M Gill B Lawlor A Lynch K S Brown P A Passmore D Craig B McGuinness S Todd C Holmes D Mann A D Smith H Beaumont D Warden G Wilcock S Love P G Kehoe N M Hooper E R Vardy J Hardy S Mead N C Fox M Rossor J Collinge W Maier F Jessen E Rüther B Schürmann R Heun H Kölsch H van den Bussche I Heuser J Kornhuber J Wiltfang M Dichgans L Frölich H Hampel J Gallacher M Hüll D Rujescu I Giegling A M Goate John Sai Keong Kauwe C Cruchaga P Nowotny J C Morris K Mayo K Sleegers K Bettens S Engelborghs P P De Deyn C Van Broeckhoven G Livingston N J Bass H Gurling A McQuillin R Gwilliam P Deloukas A Al-Chalabi C E Shaw M Tsolaki A B Singleton R Guerreiro T W Mühleisen M M Nöthen S Moebus K H Jöckel N Klopp H E Wichmann V S Pankratz S B Sando J O Aasly M Barcikowska Z K Wszolek D W Dickson N R Graff-Radford R C Petersen C M van Duijn M M Breteler M A Ikram A L DeStefano A L Fitzpatrick O Lopez L J Launer S Seshadri C Berr D Campion J Epelbaum J F Dartigues C Tzourio A Alpérovitch M Lathrop T M Feulner P Friedrich C Riehle M Krawczak S Schreiber M Mayhaus S Nicolhaus S Wagenpfeil S Steinberg H Stefansson K Stefansson J Snaedal S Björnsson P V Jonsson V Chouraki B Genier-Boley M Hiltunen H Soininen O Combarros D Zelenika M Delepine M J Bullido F Pasquier I Mateo A Frank-Garcia E Porcellini O Hanon E Coto V Alvarez P Bosco G Siciliano M Mancuso F Panza V Solfrizzi B Nacmias S Sorbi P Bossù P Piccardi B Arosio G Annoni D Seripa A Pilotto E Scarpini D Galimberti A Brice D Hannequin F Licastro L Jones P A Holmans T Jonsson M Riemenschneider K Morgan S G Younkin M J Owen M O'Donovan P Amouyel J Williams A C Naj G Jun G W Beecham L S Wang B N Vardarajan J Buros P J Gallins J D Buxbaum G P Jarvik P K Crane E B Larson T D Bird B F Boeve N R Graff-Radford P L De Jager D Evans J A Schneider M M Carrasquillo N Ertekin-Taner S G Younkin C Cruchaga John Sai Keong Kauwe P Nowotny P Kramer J Hardy M J Huentelman A J Myers M M Barmada F Y Demirci C T Baldwin R C Green E Rogaeva P St George-Hyslop S E Arnold R Barber T Beach E H Bigio J D Bowen A Boxer J R Burke N J Cairns C S Carlson R M Carney S L Carroll H C Chui D G Clark J Corneveaux C W Cotman J L Cummings C DeCarli S T DeKosky R Diaz-Arrastia M Dick D W Dickson W G Ellis K M Faber K B Fallon M R Farlow S Ferris M P Frosch D R Galasko M Ganguli M Gearing D H Geschwind B Ghetti J R Gilbert S Gilman B Giordani J D Glass J H Growdon R L Hamilton L E Harrell E Head L S Honig C M Hulette B T Hyman G A Jicha L W Jin N Johnson J Karlawish A Karydas J A Kaye R Kim E H Koo N W Kowall J J Lah A I Levey A P Lieberman O L Lopez W J Mack D C Marson F Martiniuk D C Mash E Masliah W C McCormick S M McCurry A N McDavid A C McKee M Mesulam B L Miller C A Miller J W Miller J E Parisi D P Perl E Peskind R C Petersen W W Poon J F Quinn R A Rajbhandary M Raskind B Reisberg J M Ringman E D Roberson R N Rosenberg M Sano L S Schneider W Seeley M L Shelanski M A Slifer C D Smith J A Sonnen S Spina R A Stern R E Tanzi J Q Trojanowski J C Troncoso V M Van Deerlin H V Vinters J P Vonsattel S Weintraub K A Welsh-Bohmer J Williamson R L Woltjer L B Cantwell B A Dombroski D Beekly K L Lunetta E R Martin M I Kamboh A J Saykin E M Reiman D A Bennett J C Morris T J Montine A M Goate D Blacker D W Tsuang H Hakonarson W A Kukull T M Foroud J L Haines R Mayeux M A Pericak-Vance L A Farrer G D Schellenberg John Sai Keong Kauwe C. Cruchaga C M Karch B. Sadler M. Lee K. Mayo W. Latu M. Su'a A M Fagan D M Holtzman J C Morris A M Goate L. Jones P. A. Holmans M. L. Hamshere D. Harold V. Moskvina D. Ivanov A. Pocklington R. Abraham P. Hollingworth R. Sims A. Gerrish J. S. Pahwa N. Jones A. Stretton A. R. Morgan S. Lovestone J. Powell P. Proitsi M. K. Lupton C. Brayne D. C. Rubinsztein M. Gill B. Lawlor A. Lynch K. Morgan K. S. Brown P. A. Passmore D. Craig B. McGuinness S. Todd C. Holmes D. Mann A. D. Smith S. Love P. G. Kehoe S. Mead N. Fox M. Rossor J. Collinge W. Maier F. Jessen B. Schurmann H. van den Bussche I. Heuser O. Peters J. Kornhuber J. Wiltfang M. Dichgans L. Frolich H. Hampel M. Hull D. Rujescu A. M. Goate John Sai Keong Kauwe C. Cruchaga P. Nowotny J. C. Morris K. Mayo G. Livingston N. J. Bass H. Gurling A. McQuillin R. Gwilliam P. Deloukas A. Al-Chalabi C. E. Shaw A. B. Singleton R. Guerreiro T. W. Muhleisen M. M. Nothen S. Moebus K. H. Jockel N. Klopp H. E. Wichmann E. Ruther M. M. Carrasquillo V. S. Pankratz S. G. Younkin J. Hardy M. C. O'Donovan M. J. Owen J. Williams John Sai Keong Kauwe S. Bertelsen K. Mayo C. Cruchaga R. Abraham P. Hollingworth D. Harold M. J. Owen J. Williams S. Lovestone J. C. Morris A M Goate C. Cruchaga John Sai Keong Kauwe K. Mayo N. Spiegel S. Bertelsen P. Nowotny A. R. Shah R. Abraham P. Hollingworth D. Harold M. M. Owen J. Williams S. Lovestone E. R. Peskind G. Li J. B. Leverenz D. Galasko J. C. Morris A. M. Fagan D. M. Holtzman A. M. Goate and I. The Alzheimer's Disease Neuroimaging John Sai Keong Kauwe S. Bertelsen Cruchaga K. Mayo W. Latu P. Nowotny A. L. Hinrichs A. M. Fagan D. M. Holtzman I. Alzheimer's Disease Neuroimaging A. M. Goate D. B. Clifford A. M. Fagan D. M. Holtzman J. C. Morris M. Teshome A. R. Shah John Sai Keong Kauwe D. Harold R. Abraham P. Hollingworth R. Sims A. Gerrish M. L. Hamshere J. S. Pahwa V. Moskvina K. Dowzell A. Williams N. Jones C. Thomas A. Stretton A. R. Morgan S. Lovestone J. Powell P. Proitsi M. K. Lupton C. Brayne D. C. Rubinsztein M. Gill B. Lawlor A. Lynch K. Morgan K. S. Brown P. A. Passmore D. Craig B. McGuinness S. Todd C. Holmes D. Mann A. D. Smith S. Love P. G. Kehoe J. Hardy S. Mead N. Fox M. Rossor J. Collinge W. Maier F. Jessen B. Schürmann H. van den Bussche I. Heuser J. Kornhuber J. Wiltfang M. Dichgans L. Frölich H. Hampel M. Hüll D. Rujescu A. M. Goate John Sai Keong Kauwe C. Cruchaga P. Nowotny J. C. Morris K. Mayo K. Sleegers K. Bettens S. Engelborghs P. P. De Deyn C. Van Broeckhoven G. Livingston N. J. Bass H. Gurling A. McQuillin R. Gwilliam P. Deloukas A. Al-Chalabi C. E. Shaw M. Tsolaki A. B. Singleton R. Guerreiro T. W. Mühleisen M. M. Nöthen S. Moebus K. H. Jöckel N. Klopp H. E. Wichmann M. M. Carrasquillo V. S. Pankratz S. G. Younkin P. A. Holmans M. O'Donovan M. J. Owen J. Williams C. Fenoglio D. Galimberti F. Cortini John Sai Keong Kauwe C. Cruchaga E. Venturelli C. Villa M. Serpente D. Scalabrini K. Mayo L. M. Piccio F. Clerici D. Albani C. Mariani G. Forloni N. Bresolin A. M. Goate E. Scarpini John Sai Keong Kauwe J. Wang K. Mayo J. C. Morris A. M. Fagan D. M. Holtzman A. M. Goate John Sai Keong Kauwe J. Wang S. Chakraverty A. M. Goate A. F. Henao-Martinez John Sai Keong Kauwe C. Cruchaga K. Mayo C. Fenoglio S. Bertelsen P. Nowotny D. Galimberti E. Scarpini J. C. Morris A. M. Fagan D. M. Holtzman A. M. Goate O. Mukherjee J. Wang M. Gitcho S. Chakraverty L. Taylor-Reinwald S. Shears John Sai Keong Kauwe J. Norton D. Levitch E. H. Bigio K. J. Hatanpaa C. L. White J. C. Morris N. J. Cairns A. Goate Y Hu A Hosseini John Sai Keong Kauwe J Gross N J Cairns A M Goate A M Fagan R R Townsend D M Holtzman P Nowotny X Simcock S Bertelsen A L Hinrichs John Sai Keong Kauwe K Mayo S Smemo J C Morris A Goate John Sai Keong Kauwe S. Jacquart S. Chakraverty J. Wang K. Mayo A. M. Fagan D. M. Holtzman J. C. Morris A. M. Goate P Nowotny S Bertelsen A L Hinrichs John Sai Keong Kauwe K Mayo S Jacquart J C Morris A Goate S E Wahrle A R Shah A M Fagan S Smemo John Sai Keong Kauwe A Grupe A Hinrichs K Mayo H Jiang L J Thal A M Goate D M Holtzman O Mukherjee John Sai Keong Kauwe K Mayo J C Morris A M Goate Y Li A Grupe C Rowland P Nowotny John Sai Keong Kauwe S Smemo A Hinrichs K Tacey T A Toombs S Kwok J Catanese Thomas J White Taylor J Maxwell P Hollingworth R Abraham D C Rubinsztein C Brayne F Wavrant-De Vrièze J Hardy M O'Donovan S Lovestone J C Morris L J Thal M Owen J Williams A Goate O Mukherjee P Pastor N J Cairns S Chakraverty John Sai Keong Kauwe S Shears M I Behrens J Budde A L Hinrichs J Norton D Levitch L Taylor-Reinwald M Gitcho P H Tu L Tenenholz Grinberg R M Liscic J Armendariz J C Morris A M Goate A Grupe Y Li C Rowland P Nowotny A L Hinrichs S Smemo John Sai Keong Kauwe T A Maxwell S Cherny L Doil K Tacey R van Luchene A Myers F Wavrant-De Vrièze M Kaleem P Hollingworth L Jehu C Foy N Archer G Hamilton P Holmans C M Morris J Catanese J Sninsky T J White J Powell J Hardy M O'Donovan S Lovestone L Jones J C Morris L Thal M Owen J Williams A Goate S Smemo P Nowotny A L Hinrichs John Sai Keong Kauwe S Cherny K Erickson A J Myers M Kaleem L Marlowe A M Gibson P Hollingworth M C O'Donovan C M Morris P Holmans S Lovestone J C Morris L Thal Y Li A Grupe J Hardy M J Owen J Williams A Goate B K Suarez C Taylor S Bertelsen L J Bierut G Dunn C H Jin John Sai Keong Kauwe A D Paterson A L Hinrichs G Dunn A L Hinrichs S Bertelsen C H Jin John Sai Keong Kauwe B K Suarez L J Bierut A L Hinrichs S Bertelsen L J Bierut G Dunn C H Jin John Sai Keong Kauwe B K Suarez John Sai Keong Kauwe S Bertelsen L J Bierut G Dunn A L Hinrichs C H Jin B K Suarez P Nowotny A L Hinrichs S Smemo John Sai Keong Kauwe T Maxwell P Holmans M Hamshere D Turic L Jehu P Hollingworth P Moore L Bryden A Myers L M Doil K M Tacey A M Gibson I G McKeith R H Perry C M Morris L Thal J C Morris M C O'Donovan S Lovestone A Grupe J Hardy M J Owen J Williams A Goate Y Li P Nowotny P Holmans S Smemo John Sai Keong Kauwe A L Hinrichs K Tacey L Doil R van Luchene V Garcia C Rowland S Schrodi D Leong G Gogic J Chan A Cravchik D Ross K Lau S Kwok S Y Chang J Catanese J Sninsky White T J Hardy J Powell S Lovestone J C Morris L Thal M Owen J Williams A Goate A Grupe John Sai Keong Kauwe Dennis K Shiozawa Richard Paul Evans
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